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Athyrium crenulatoserrulatum Makino

Aspleniaceae · Khabarovsk, Primorye +5 · listed by its genus as a fern or moss

English names not checked: Wikidata's common names were not read for this page.

Photographs not checked iNaturalist and GBIF media were skipped (not asked) when this page was built. Not asked, or not answered, is not "none": nothing here claims there is none. Why the site says it this way.
Add one to my plants
Sow seed
Habitat figures
Cold floor (1 night in 100)
-19.5°C
Record low -30.1 °C in 44 years, at a typical spot in the range NASA POWER
Warmest month, mean daily high
25°C
Aug at the habitat CHELSA
Rain a year (median across the range)
1,612mm
12 months of 25 mm or more in the median year CHELSA
Open-sky light
15–36DLI
Lowest to highest month (DLI) CHELSA
The year in your months, northern hemisphere
  1. Jan
  2. Feb
  3. Mar
  4. Apr
  5. May
  6. Jun
  7. Jul
  8. Aug
  9. Sep
  10. Oct
  11. Nov
  12. Dec
Wet summers: the rain comes May to September, read as a summer growing season. rain rule, CHELSA
Northern hemisphere unless you set your site. The chart is in the habitat's months.

About the genus · Athyrium

Athyrium (lady-fern) is a genus of about 180 species of terrestrial ferns, with a cosmopolitan distribution. It is placed in the family Athyriaceae, in the order Polypodiales. Its genus name is from Greek a- ('without') and Latinized Greek thyreos ('shield'), describing its inconspicuous indusium . More on Wikipedia ›

Quoted from Wikipedia, "Athyrium", CC BY-SA 4.0.

Cultivation

Grouped as a fern or moss, listed under the genus Athyrium in the archetype table, and Kew describes it as “perennial”.

Seasons

The rain rule reads a summer growing season: 70% of the median year's rain (1,069 mm of 1,460 mm) falls in May to September, warmer than the year (wet-season mean 18.6 °C against a yearly mean of 9.1 °C). Months are the habitat's, northern hemisphere; shifted six months for a southern-hemisphere collection: November to March.

Rain

1,460 mm in the median year at the habitat, 1,069 mm of it May to September. Wettest month July at 295 mm, driest month December at 30 mm. Across the grid cells of the range the year's total runs 894 mm to 2,091 mm, the 10th to 90th percentile of each cell's own year. The median of the cells' own yearly totals, 1,612 mm, is the year's rain at the top of the page; the 1,460 mm here is the median year's, the twelve monthly medians added, which the rain rule reads. Habitat calendar, northern hemisphere; median year across the grid cells of the range, CHELSA.

Light

Open sky over the habitat: 15 to 36 mol/m²/day across the year (daily light integral, median year across the grid cells of the range, CHELSA; the lowest 10th-percentile month across the cells is 11, the highest 90th-percentile month 37).

Warmth and air
Temperature

Coldest: the 1st-percentile night over 44 years at a typical spot in the range is -19.5 °C, the absolute minimum -30.1 °C, 6,728 frost nights in 44 years, 152.9 a year (NASA POWER daily minima). Warmest: the 99th-percentile day is 30.0 °C. Monthly means: the coldest month's mean nightly low -11.5 °C in January (across the grid cells of the range -21.7 °C to -2.1 °C), the warmest month's mean daily high 25.1 °C in August (median year across the grid cells of the range, CHELSA). Cold floor: -19.5 °C, which is the 1st-percentile night over 44 years at a typical spot in the range (NASA POWER).

Humidity

Relative humidity at the habitat: 55 to 69% across the year (monthly means, median year across the grid cells of the range, CHELSA). A figure about the air, saying nothing about how the plant takes water.

How this section is made

Each card states this species' habitat figures with their sources, and what two fixed rules read from them; nothing here says what the plant does, wants or tolerates. The rules · the terms.

Seasons. Read from the median year of CHELSA monthly rainfall and mean temperature across the grid cells of the range: the rain rule takes the smallest set of months carrying 70% of the rain and calls it winter when its mean is more than half a degree below the year's, summer when more than that above, and neither in between; the temperature rule, used only under 120 mm, names the cooler six months when they are at least a degree cooler than the other six. Months are shifted for the other hemisphere only where the temperature curve moves 4 °C or more and the habitat lies 10° or more from the equator. A reading of the curves, not an observation of the plant.

Rain. CHELSA monthly precipitation, median year across the grid cells of the range. The rain that falls where the species is recorded; nothing about how the plant takes water.

Light. CHELSA shortwave radiation at each envelope cell, a daily-mean flux taken to a daily total and converted at 2.07 mol of PAR per MJ. The sky over the habitat, measured; what reaches a plant under a rock, a shrub or a shade cloth is not.

Temperature. NASA POWER daily minima and maxima over 44 years at a typical spot in the range, lapse-corrected +222 m to its elevation; CHELSA monthly means, median year across the grid cells of the range. The cold floor is the 1st-percentile night named in its sentence. Not a measured survival limit for any plant in a pot.

Humidity. CHELSA relative humidity, median year across the grid cells of the range.

Group. Listed under the genus Athyrium in the archetype table, and Kew describes it as “perennial”. A group is taken only where Kew's description of the species agrees with the table's listing. The table gives no figure and no prose; the cold floor is the habitat's alone.

Climate across the habitat

Habitat months, northern hemisphere

Habitat climate of Athyrium crenulatoserrulatum through the yearWarmest month August, mean daily high 25 °C; coldest month January, mean nightly low -11.5 °C; 1,460 mm of rain in the median year (the twelve monthly medians added). Medians across the range, from CHELSA; the extremes at the edge from NASA POWER. Habitat months, northern hemisphere.-40°-20°0°20°40°frost30.0° 99th-percentile day, NASA POWER-30.1° lowest night in 44 years, NASA POWER°C · day and nightcold quarter0100200300400500mm rainDLI 15–36RH 55–69%JanFebMarAprMayJunJulAugSepOctNovDec
day night rain medians and 10th–90th percentile across 137 habitat cells, CHELSA cold quarter: the three months around the coldest month's mean nightly low extremes over 44 years at a typical spot in the range, from NASA POWER, marked at the edge, in no month DLI, mol/m²/dayRH %each on its own scale
Figures by month
JanFebMarAprMayJunJulAugSepOctNovDec
Day °C-1 / -10–41 / -6–66 / 0–1013 / 8–1718 / 14–2121 / 17–2524 / 21–2725 / 22–2821 / 18–2415 / 11–198 / 1–131 / -7–7
Night °C-12 / -22–-2-9 / -18–-2-4 / -10–23 / -1–78 / 5–1213 / 11–1718 / 15–2118 / 16–2213 / 10–186 / 2–12-1 / -8–6-7 / -18–1
Rain mm33 / 15–8341 / 16–9167 / 32–14392 / 52–147119 / 85–174170 / 112–237295 / 161–463283 / 158–413202 / 120–29370 / 53–18158 / 36–11430 / 21–74
DLI17 / 13–1922 / 19–2429 / 26–3034 / 33–3536 / 34–3734 / 30–3528 / 26–3228 / 27–3226 / 23–2723 / 19–2417 / 14–1915 / 11–17
RH %58 / 54–6757 / 54–6456 / 54–6055 / 53–6058 / 56–6362 / 60–6969 / 65–7267 / 64–7063 / 61–6758 / 57–6358 / 54–6258 / 54–65
How this section is made
Each figure is the median across the 137 grid cells holding the 161 in-range records, with the 10th–90th percentile span across those cells after the slash where it differs. A dash is a month one or more of those cells has no figure for in the grid (a variable CHELSA does not carry there), so no median is taken rather than one over fewer cells. Extremes and elevation were read at a typical spot in the range, cell 1055:6159 (37.225, 127.975). Over 44 years there: absolute minimum -30.1 °C, 1st-percentile night -19.5 °C, 99th-percentile day 30.0 °C. Normals: CHELSA V2.1 1981–2010 climatology, 0.05° cells (each the mean of ~36 1 km pixels). Across the range: median and 10th–90th percentile of each month across the 137 distinct grid cells holding 161 of the 161 in-range records; extremes and elevation at the typical cell 1055:6159 (its coldest month's mean nightly low nearest the median across cells); its position is the cell centre. Extremes: NASA POWER (MERRA-2) daily 1981–2024, cell 37.00_128.125, lapse-corrected +222 m at 6.5 °C/km. Elevation: ETOPO 2022, 589 m (cell mean).

Natural habitat

Native range as published by WCVP; the marker is where the records are densest and decides nothing: the climate was read across the cells of the in-range records placed well enough to read one, not at the marker.
Openly licensed records inside the range, framed on where they fall.

This species has no GBIF key of its own. Its records are GBIF's under the key of the name Kew's World Checklist holds as this plant: 7228804 (Cornopteris crenulatoserrulata (Makino) Nakai, 1999 records). No name was matched to GBIF; the keys are read from Kew's synonymy. How.

NativeKhabarovsk, Primorye, Manchuria, China North-Central, China Southeast, Japan, Korea · regions are the level-3 units of the World Geographical Scheme for Recording Plant Distributions, as Kew's World Checklist records ranges, some under historical names such as Yugoslavia or Czechoslovakia (how).
Map marker36.28, 138.597 · in the densest population, 986 records (67% of those in range)
Kew's description“perennial” · “temperate” · quoted from WCVP, RBG Kew (CC BY 4.0); not derived here
Evidence usedThe map marker rests on all 1461 georeferenced records inside the native range; the climate across the range on the 161 of them placed to within 10 km; the map shows only the 1354 openly licensed ones, which alone would put the marker in the same place. 1 record outside the WCVP native range not used. 1300 in-range records are placed to worse than 10 km, or with no stated accuracy (admitted only as a person's observation to three decimals), and stay on the map but off the climate.
Other GBIF keysRecords also from GBIF's key 7228804 (Cornopteris crenulatoserrulata (Makino) Nakai): Kew's file holds it as Cornopteris crenulatoserrulata (Makino) Nakai (synonym under Athyrium crenulatoserrulatum Makino); 1,999 records, from GBIF's download.
How this section is made
The map marker: the openly licensed record nearest the middle of the densest 1° population inside the densest 2° block of the 1461 in-range records; it decides nothing. Records outside the native range, living specimens and records marked introduced are not used; which records count.

Photographs

Not checked. iNaturalist and GBIF media were skipped (not asked) when this page was built. Not a statement that none exist.

Similar habitat climate The 6 nearest by mean daily high, mean nightly low and rain, month by month. How.

Adenocaulon himalaicum Adenocaulon himalaicum Asian trailplant Photo: CC BY, W RaoFilipendula glaberrima Filipendula glaberrima Korean meadowsweet Photo: CC BY, Tatyana PetrenkoActaea asiatica Actaea asiatica Asian baneberry Photo: CC BY, Jane CharlenDryopteris crassirhizoma Dryopteris crassirhizoma Shield fern Photo: CC BY, Violetta DzizyurovaChrysosplenium pilosum Chrysosplenium pilosum Hairy golden saxifrage Photo: CC BY, Tatyana PetrenkoCorylus sieboldiana Corylus sieboldiana Japanese hazel Photo: CC BY, Станислав Чуванов

Other Athyrium 13 in the reference

Athyrium americanum Athyrium americanum Alpine ladyfern Photo: CC BY, James H. ThomasAthyrium angustum Athyrium angustum Subarctic lady fern Photo: CC BY, Karen SaundersAthyrium asplenioides Athyrium asplenioides Southern lady fern Photo: CC BY, David WilkinsAthyrium cyclosorum Athyrium cyclosorum Western Lady Fern Photo: CC0, Ellyne GeurtsAthyrium distentifolium Athyrium distentifolium Aspleniaceae Photo: CC BY-SA, Jerzy OpiołaAthyrium filix-femina Athyrium filix-femina Lady-fern Photo: CC0, author not stated Athyrium iseanum Narrow-leaf ladyfern Athyrium microphyllum Athyrium microphyllum Akolea Photo: CC0, author not statedAthyrium otophorum Athyrium otophorum Eared lady fern Photo: CC0, author not statedAthyrium rubripes Athyrium rubripes Athyriaceae Photo: CC BY, Violetta DzizyurovaAthyrium strigillosum Athyrium strigillosum Athyriaceae Photo: CC BY, Jacy ChenAthyrium vidalii Athyrium vidalii Vidal’s ladyfern Photo: CC BY-SA, harum.koh all 14 ›

Names & registers

Also known as Cornopteris coreana; Dryopteris austroussuriensis; Dryopteris crenulatoserrulata; Neoathyrium crenulatoserrulatum; Neoathyrium crenulatoserrulatum var. coreanum and 7 more. (WCVP, RBG Kew)

IPNIPOWOWikidataWFO

The record

Family
Aspleniaceae
Described by
Makino
Native to
Khabarovsk, Primorye, Manchuria +4
Wild records
1461 in range · 1354 open

Every figure here is derived from public data by a stated rule and names its source; nothing about this species is written for its page by a person or by AI, apart from the marked, credited quotations. How →

Where this page came from built 2026-10-22
Kew's names (WCVP) kew.names answeredas of 2026-06-04Kew's entry Athyrium crenulatoserrulatum Makino (plant_name_id 3173721), read from Kew's names file
GBIF species record gbif.species not askedGBIF's backbone holds no one accepted species of this name, and a name it holds only as another species' synonym is not taken for this one, so GBIF's records were not asked
GBIF synonyms gbif.synonyms not askedGBIF's backbone holds no one accepted species of this name, and a name it holds only as another species' synonym is not taken for this one, so GBIF's records were not asked
GBIF common names gbif.vernacular not askedGBIF's backbone holds no one accepted species of this name, and a name it holds only as another species' synonym is not taken for this one, so GBIF's records were not asked
WCVP native range wcvp.distribution answeredas of 2026-06-04
GBIF occurrence records gbif.occurrences answeredas of 2026-10-21this species has no GBIF key of its own; its records are read under GBIF's keys for the names Kew's file holds as this plant, never by a name matched; GBIF occurrence download https://doi.org/10.15468/dl.hbdw64, of 2026-10-21; 1999 records of GBIF's key 7228804, Cornopteris crenulatoserrulata (Makino) Nakai (Kew's file holds it as Cornopteris crenulatoserrulata (Makino) Nakai (synonym under Athyrium crenulatoserrulatum Makino)): the GBIF download; 14 records within 1 km of a botanic garden, herbarium or zoo left out (11 at a botanic garden or arboretum, 3 at a zoo; Wikidata's list)
Habitat climate climate answeredasked 2026-10-22
Daily extremes (NASA POWER) climate.extremes answeredasked 2026-10-22NASA POWER (MERRA-2) daily 1981–2024, cell 37.00_128.125, lapse-corrected +222 m at 6.5 °C/km
Wikidata wikidata answeredasked 2026-10-22found by Kew's IPNI identifier 17054650-1, POWO 17054650-1
Wikipedia wikipedia nothing to reportasked 2026-10-22
iNaturalist photographs, wild inat.photos.wild not asked
iNaturalist photographs, cultivated inat.photos.cultivated not asked
GBIF media gbif.media not askedGBIF's backbone holds no one accepted species of this name, and a name it holds only as another species' synonym is not taken for this one, so GBIF's records were not asked
Wikimedia Commons commons nothing to reportasked 2026-10-22
OpenAlex papers openalex not askednot yet asked for this reference; a later fill takes the papers from OpenAlex's snapshot

Every source is recorded as having answered, reported nothing, refused, failed, not been asked, or answered twice where one answer is taken (“not taken: two items answered”), and, for a source asked when this page was built, when: the figures are as the source held them that day; a source read from a file (Kew's World Checklist, a GBIF download) says "as of" the file's day instead; a row carried from an earlier build of the page names that build instead. Only “nothing to report” is ever shown as an absence. Records came from 66 datasets via GBIF.org; each carries its own licence.

Cultifolio

Opening your plants…